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Yimmit....Keep in mind that skulls only activate when they are picked up while playing LEGENDARY, and their effects are only working while playing on LEGENDARY.
Once you have the skull (and the text of the name appears on the screen) it is yours as are its effects, you can drop it, carry it, beat something with it or just plain stand there. There is only one skull you can keep as a third weapon (for melee only), I think it is Angry, but I am not sure. You do not have to complete the level or anything. Normally when I put H2 in, I go get sputnik, before the noise has even stopped I have saved and quit to load another level. Usually Delta Halo (I LOVE THAT LEVEL) to go get envy, then as soon as I have the skull, its save and quit and load another level, usually outskirts or cairo station. From then on I play the campaign in order and pick up skulls as I go.
Catch is one of the most fun (causes each killed enemie to drop two grenades, and throw more at you), but watch out, with catch and sputnik on at the same time it is pretty hairy not only dodging all the extra (and well aimed) covie plasmas, but there are alot of unexploded grenades lying around to create some huge explosions with. Those can be just as dangerous to you as they are to others!
Just remember that until you either eject the disk, change profiles, or turn off your box, the skull effect is active.