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I do not think that the Keyes flood is a pre-cursor to the Gravemind flood. It seems that the Gravemind has been in Delta Halo for a loooooooooooooong time. The Keyes flood was just another cool thing in the game, a flood not built for combat, but for piloting the covie ship off the ring so the flood could escape. It is more than likely to have gone like this:
Bungie Employee #1-"Wouldnt it be cool if Keyes got turned into a flood!"
Bungie Employee #2-"Yeah, that would be sweet, but it cant be a combat flood, or a carrier, and we cant make him a popcorn!"
Bungie Employee #3-"You guys are idiots, he is a pilot, he has to be a flying flood!"
Bungie Employees #1 & #2 in unison-"YOU ARE A MORON #3!"
GOD-"Hey, IDIOTS, make a new flood, one that is cool, different and seen no where else in the game, and if I am pleased, I will not SMITE YOU!"
I am pretty sure that is exactly how it got in the game!